When Sammy and Ray find themselves trapped in The Tank, an underwater aquarium, they must hatch a plan to escape and reunite with their grandkids, Ricky and Ella, and protect them from the dangers of the ocean.

Born on a Baja, California beach in 1959, new hatchling Sammy races across the beach to the ocean while avoiding being caught by seagulls and crabs. Thus begins Sammy's incredible fifty-year ocean journey where he overcomes obstacles, both natural and man-made, while trying to fulfill his dream of travelling around the world. Along the way he meets his best friend, a fellow turtle named Ray, and never forgets about Shelly, a turtle he saved that first day on the beach and the one he's always loved.